The Harlequin Ladybird Survey
Harlequin ladybird elytra
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Harlequin Ladybird Project Publications

Brown, P.M.J., Roy, H.E., Ware, R.L. and Majerus, M.E.N. (2006) Distribution and identification of the harlequin ladybird Harmonia axyridis. Atropos 28: 63-69.

Brown, P.M.J. & Roy, H.E. (in press) The spread of the Harlequin Ladybird Harmonia axyridis in late 2006. Atropos.

Brown, P.M.J., Roy, H.E. and Majerus, M.E.N. (2006) Ladybird recording in Cambridgeshire and Huntingdonshire. Nature in Cambridgeshire 48: 12-16.

Brown, P.M.J., Roy, H.E. and Majerus, M.E.N. (in press) The spread of the harlequin ladybird in Cambridgeshire and Huntingdonshire. Nature in Cambridgeshire.

Brown, P.M.J., Roy, H.E. and Majerus, M.E.N. (2007) Update on the spread of an invasive ladybird. Biocontrol News and Information 28(1): 16-17.

CAB International (2006) Harmonia axyridis [original text by H.E. Roy, P.M.J. Brown and M.E.N. Majerus]. Crop Protection Compendium, 2006 edition. Wallingford, UK: CAB International.

Majerus, M.E.N. and Roy, H.E. (2005) Scientific opportunities presented by the arrival of the Harlequin ladybird, Harmonia axyridis, in Britain. Antenna 29(3): 196-208.

Majerus, M.E.N., Mabbott, P., Rowland, F. and Roy, H.E. (2006) The harlequin ladybird, Harmonia axyridis, arrives in Britain. Entomologist’s Monthly Magazine 142: 87-92.

Majerus, M.E.N., Strawson, V. and Roy, H.E. (2006) The potential impacts of the arrival of the Harlequin ladybird, Harmonia axyridis (Pallas) (Coleoptera: Coccinellidae), in Britain. Ecological Entomology 31(3): 207-215.

Majerus, M.E.N. and Roy, H.E. (2005) Arrival of the Harlequin Ladybird in Britain. Biological Sciences Review, Sept. 2005: 29-32.

Majerus, M.E.N., Roy, H.E., Brown, P.M.J., Ware, R.L. and Shields, C. (2006) A Guide to the Ladybirds of the British Isles (fold-out chart). Field Studies Council, Shrewsbury.

Roy, H.E., Brown, P.M.J., James, T.J., Munford, J. and Majerus, M.E.N. (2005) Monitoring an Alien: Harmonia axyridis. Journal of Practical Ecology and Conservation Special Series, No.4: 77-82.

Roy, H.E., Rowland, F., Brown, P.M.J., and Majerus, M.E.N. (2005) Ecology of the harlequin ladybird: a new invasive species. British Wildlife 16(6): 403-407.

Roy, H.E. and Majerus, M.E.N. (2005) Unlovely Ladybirds? Biological Sciences Review, 17(4): 42.

Roy, H.E., Brown, P.M.J. and Majerus, M.E.N. (2006) Harmonia axyridis: A successful biocontrol agent or an invasive threat? In: An ecological and societal approach to biological control (eds.: J. Eilenberg & H. Hokkanen). © Kluwer Academic Publishers, Netherlands.

Ware, R.L., Majerus, M.E.N., Roy, H.E. and Symington, F. (2005). The harlequin ladybird arrives in Britain: a threat to our native species? Bulletin of the Amateur Entomologists' Society 64: 175-186.

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